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After crushing and the appropriate cleaning steps (Ren et al., 2012; Ren et al., 2015), shell-bound N was released by dissolution with hydrochloric acid, and organic N converted to nitrate using a basic persulfate oxidising reagent (POR; a potassium persulfate/sodium hydroxide solution) (Nydahl, 1978; Knapp et al., 2005). The \"denitrifier method\"\u00a0was used to convert sample nitrate to N<sub>2</sub>O gas for isotope analysis (Sigman et al., 2001; Casciotti et al., 2002; Weigand et al., 2016). See methods section of Smart et al. (2018) for more details of sample collection, isotope analysis and data quality.</p>\n<p>The isotopic composition of N<sub>2</sub>O was measured at Princeton University by GC-IRMS using a Thermo MAT 253 mass spectrometer with a purpose-built on-line N<sub>2</sub>O extraction and purification system. See methods section of Smart et al. 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